Each chapter has its own title and focus "Hulkamania Runs Wild" focuses on Hulk Hogan and early WrestleManias, while "The New Generation" showcases the rise of Bret Hart and Shawn Michaels in the mid-1990s. In 45 primary matches, the player competes through storylines of various lengths, switching predetermined characters as they are completed. Sometimes, the player must follow a guided sequence of inputs, rather than freely control the character. In Wrestlemania mode, matches have primary objectives, required to beat the game, and bonus historical objectives, which result in recreated WrestleMania moments. The mode features a distinct roster of wrestlers and alternate gimmicks, who wrestled at WrestleMania (especially in main events). Like WWE Legends of WrestleMania and WWE '13, the campaign revolves around historical stories rather than original ones.
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WWE 2K14 features a new single player campaign called "The 30 Years of WrestleMania", a nostalgia-based gameplay mode that recreates nearly three decades of WWE history, including 45 matches, classic WWE footage and many legendary characters. The ability to create a custom WWE championship is also available, having been made more reminiscent to previous iterations after having limited creativity in WWE '13. The player continues to have the option to create their own entrances and finishers. While the mode itself remains much of the same, the ability to use existing superstars as a template for new custom variants has been added. The long-standing mode, "Create-a-Superstar" has been tweaked to have an expanded character save slot that can be filled with up to 100 created superstars. Few new matches have been added, though a Slobberknocker Mode, adapted from The Streak season mode, had been added as a gauntlet match. The nearfall system too has been improved, with more two counts. Seven new OMG moments have been added, allowing certain finishers to be done on two opponents, and players to interact with the arena environment. Some characters can launch an opponent into the air and catch them for a catch finisher. Catapult finishers can now be used offensively. All chain reversals now result in offensive attacks, to make for quicker matches by avoiding continuous reversal stalemates. Before running, characters display "starting up" animations, to prevent players from spamming running attacks. Characters also move faster than previously. These include the navigation system, allowing for more fluid walking, running and dragging motions.
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Many of the gameplay features from WWE '13 have been updated.